Author: (Sym.) de Wit
Morphological description
Tree up to 30(-39) m high and 0.4(-1,8) m in diameter.
Leaves (2-)4(-6)-foliolate; stipules auriculate to subfalcate, up to 1.5 by 1 cm; petiole and rachis 4.5-12 cm, glabrous. Leaflets coriaceous, elliptic-oblong, often unequal-sided, 5-9(-14) by 3.5-6(-8) cm; apex acute or shortly acuminate; base rotund; glossy, pellucid-dotted, glabrous except sometimes sparsely hairy on the midrib or at the base on the lower surface, glabrescent; nerves many, distinct or visible on both surfaces; often with 1-3 elliptic, slightly depressed glands towards the basal part of the margin; petiolules 4-7 mm (palustris Copaifera lv/pod FM24 ).
Inflorescences 4-11 cm long; pedicels 3-4 mm.
Flower. Calyx lobes elliptic, 4-7.5 by 2-4 mm, puberulous outside, densely tomentose inside. Stamens alternately long and short; filaments 7-10 mm; anthers c. 2 mm long. Ovary 2-3 by 1-1.5 mm, style 4.5-5 mm.
Pods 4.5-7.5 by 3.5-4 cm (palustris Copaifera lv/pod FM24 ).
Seeds 2.4-2.7 by c. 1.5 cm, glossy brown or black; aril narrowly 2-lobed, enclosing the seed.
Distribution
Malesia: Borneo (Brunei, Sabah, Sarawak, Kalimantan).
Habitat & Ecology
In freshwater peat swamp forest near the coast, sometimes also found in dry lowland forest, once found in secondary heath forest, 0-30 m altitude Fl. April, October; fr. July.
Uses
The timber of the present species is similar to that of Sindora species and can be used for the same purposes (Sambas in Soerianegara & Lemmens (eds.), Pl. Res. SE Asia (PROSEA Handb.) 5 (1), Major commercial timbers (1993) 437.