Bauhinia kockiana

Author: Korthals

A very polymorphic species; see also Note 2 under Bauhinia finlaysoniana Larsen & Larsen (1996: 522) have also here recognized a few distinctive varieties. It must be admitted that the delimitation towards B. finlaysoniana is not always clear; in spite of this Larsen & Larsen (1996) have maintained the two with specific status.
(see also description of Genus Bauhinia ; Subgenus Phanera and Section Palmatifolia )

KEY TO THE VARIETIES
1a.   Filament of fertile stamens short, c. 3 mm, hairy as the staminodes.
................. Bauhinia kockiana beccarii
1b.   Filament of fertile stamens longer, 6-13 mm, glabrous as the staminodes.
................. 2.
2a.   Pedicels short, 1-1.5 cm; hypanthium 2-2.7 cm long; bracts large, 8-10 by 4 mm.
.......... Bauhinia kockiana brevipedicellata
2b.   Not this combination...................... 3.
3a.   Hypanthium short, 4-8 mm; leaves rigidly coriaceous, small, 3.5 by 1.3 to 9 by 3 cm, glabrous.
........................ Bauhinia kockiana calcicola
3b.   Hypanthium and leaves otherwise.
................................................... 4.
4a.   Ovary silky, light brown, appressed hairy all over, hairs retained after anthesis; leaves lanceolate, 11.5 by 2.5 to 20 by 5 cm.
................. Bauhinia kockiana angustifolia
4b.   Ovary hairy along sutures only, occasionally hairy all over but after anthesis laterally glabrous...................... 5.
5a.   Leaves small, 4 by 2 to 8 by 3.5 cm, densely rusty hairy at nerves beneath, petioles and inflorescences.
.............. Bauhinia kockiana sericeinervia
5b.   Not this combination......................... 6.
6a.   Petioles thick, densely hairy; lower surface of leaves of young shoots and inflorescences densely hairy.................... 7.
6b.   Not this combination......................... 8.
7a.   Leaves sub-5-nerved (= with 3 long nerves reaching apex, and 2 short, distinct, basal nerves) base ± cordate, rarely obtuse, persistently darkbrown, densely hairy beneath; pedicels slender, patently hairy.
....................... Bauhinia kockiana velutina
7b.   Leaves 3-nerved to sub-5-nerved, base cuneate to obtuse, reddish brown beneath or mainly on the nerves; pedicels velvety hairy to puberulous.
................... Bauhinia kockiana scarlatina
8a.   Leaves rigidly coriaceous, large, 16 by 5 to 21 by 7.5 cm, glabrous, apex acuminate, base ± truncate.
..................... Bauhinia kockiana bakoensis
8b.   Leaves chartaceous to coriaceous, varying in form, size and hairiness, base cuneate to ± truncate.
..................... Bauhinia kockiana kockiana

a. Bauhinia kockiana Korth. var. kockiana

Morphological description
Large, tendrilled liana with flattened stems which may reach up to at least 30 m; young shoots ± hairy, usually reddish-brown tomentose, later glabrous (kockiana Bauh in tree 431668).

Leaves (kockiana Bauhinia infl 431626) entire, glabrous or sparsely hairy when young, shining on upper surface; stipules auriculate, puberulous, 4-7 mm long, early caducous; petiole usually short, c. 5 mm, reddish brown puberulous; lamina chartaceous to coriaceous, 7-13 by 4-6 cm, narrowly ovate, elliptic to oblong, apex acuminate, often with a slightly emarginate tip, usually 3-nerved, sometimes sub-5-nerved, base obtuse.

Inflorescence (kockiana Bauh infl 431667) lateral and terminal corymbs or short racemes, often broader than long, varying in hairiness but in early stages and the axis usually reddish or brownish tomentose; pedicels 4-6(-7) cm; bracts ovate-oblong, c. 1 cm, caducous; bracteoles linear, few mm long, inserted below the middle.

Flowers: (kockiana Bauh fl 431667) Buds globose, apiculate, ± prominently longitudinally ridged, 3-5 mm diameter. Hypanthium narrow tubular, finely striate, 11-18 mm long, 1-2 mm diameter, only slightly dilated at the base. Calyx splitting into 2-3 segments, glabrous inside. Petals yellow to orange turning red, short-clawed, broadly obovate with crenulate, undulate or flat margin, 2-2.5 cm long; outside hairy, inside almost glabrous. Fertile stamens 3; filaments glabrous, up to 8 mm long; anthers small, ellipsoid, 2-3 mm, opening by a central pore in each theca; reduced stamens and staminodes 3-5. Ovary shortly stipitate, 5-9 mm, brownish or reddish hairy, at least along the sutures; style c. 6 mm, glabrous or nearly so; stigma capitate.

Pods narrowly elliptic, glabrous, c. 15 cm long.

Seeds flat, broadly ovate, c. 2 cm across.

Distribution
Malesia: widely distributed from the Malay Peninsula (not reaching Thailand) throughout Borneo to the Lesser Sunda Islands; not reaching the Philippines.

Habitat
From sea level up to 1200 m, in a variety of habitats.

Vernacular names
Tagulogulog, Urat tembadau, Tali adap, Tongkurangan (N Borneo), Djemprangan (E Borneo), Tagalap (Dusun Kina-batabgab); Akar djangan (Sumatra, Jambi).

Uses
According to de Wit (1956: 521), in Sabah used in religious ceremonies.

b. Bauhinia kockiana Korth. var. angustifolia K. & S.S. Larsen

Morphological description
Deviates from the other varieties by the leaves being lanceolate, 11.5-20 by 2.5-5 cm, and the ovary light brown silky appressed-hairy all over, the indumentum retained after anthesis.

Distribution
Borneo, endemic to Sabah.

Habitat & Ecology
On ultrabasic soil.

c. Bauhinia kockiana Korth. var. bakoensis K. & S.S. Larsen

Morphological description
Leaves large, ovate-oblong to lanceolate, glabrous, strongly coriaceous, 16-21 by 5-7.5 cm, apex acuminate, base ± truncate; nerves 3, sometimes with 2 short basal ones; petiole 1-2 cm.

Distribution
Borneo: Sarawak, only known from Bako National Park.

Habitat
Heath forest on humid podsol soil on sandstone, from sea level up to 600 m altitude.

d. Bauhinia kockiana Korth. var. beccarii K. & S.S. Larsen

Morphological description
Deviates from the other varieties by the filaments of the fertile stamens being short, c. 3 mm, and hairy as the staminodes.

Distribution
Borneo (Sabah).

Habitat
Primary forest, 600 m altitude.

e. Bauhinia kockiana Korth. var. brevipedicellata K. & S.S. Larsen

Morphological description
Deviates from the other varieties by its short pedicel, 1-1.5 cm long, the long hypanthium, 2-2.7 cm, the large, broad bracts, 8-10 by 4 cm, and the bracteoles 4 by 1 mm.

Distribution
Borneo, only known from the type collected on Bukit Raya, C Kalimantan.

Habitat
Primary forest.

f. Bauhinia cockiana Korth. var. calcicola K. & S.S. Larsen

Morphological description
Leaves rigidly coriceous, narrowly elliptic, glabrous, small, 3.5-9 by 1.3-3 cm, hypanthium short, 4-8 mm.

Distribution
Borneo (Sarawak, only known from Bau distr.)

Habitat
On limestone at low altitude.

g. Bauhinia cockiana Korth. var. scarlatina (Camm.) K. & S.S. Larsen

Morphological description (kockiana scar Bauh compl 139508)
This variety has larger leaves , (7-)10-17 by 3-6 cm, larger on the basal shoots; 3- to sub-5-nerved, base cuneate to rounded, reddish-brown hairy beneath, mainly on the nerves; pedicels velvety hairy to sparsely pubescent. Flowers rather large.

Distribution
Borneo (Kalimantan).

Habitat
Forest, up to 700 m altitude.

h. Bauhinia kockiana Korth. var. sericeinervia (de Wit) K. & S.S. Larsen

Morphological description
Characterized by small leaves, from 4 by 2 to 8 by 3.5 cm, densely rusty hairy at nerves beneath and at petioles and inflorescences. Flowers somewhat smaller than in var. kockiana.

Distribution
Sumatra.

Habitat
In primary forest at lower altitudes.

i. Bauhinia kockiana Korth. var. velutina (de Wit) K. & S.S. Larsen

Morphological description
Characterized by leaves sub-5-nerved, crossveins prominent, base ± cordate, rarely obtuse, persistently dark brown, densely hairy beneath; pedicels slender, patently hairy.

Distribution
Borneo, from W Kalimantan to Sarawak.

Habitat
Riverine forests, altitude 150 m.

Note
Only known from few collections.

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